Biodiversity of Venezuela: Four new species of Tityus

I've learned about another new paper from the late Manuel Gonzalez Sponga. Here four new species of Tityus Koch, 1836 (Buthidae) are described from the Venezuelan states of Monagas, Sucre and Bolivar:

Tityus longidigitus Gonzalez Sponga, 2008
Tityus quiriquirensis Gonzalez Sponga, 2008
Tityus romeroi Gonzalez Sponga, 2008
Tityus sanfernandoi Gonzalez Sponga, 2008

Abstract:
Four new species of Venezuelan scorpions of the genus Tityus Koch. 1836 (Buthidae): Tityus longidigitus, Tityus quiriquirensis, Tityus romeroi and Tityus sanfernandoi are described. They live in the oriental and Imataca mountain range regions in Venezuela. in altitudes between 100 and 200 m. Drawing of significant morphological characters are presented to diagnose the species: tables with morphometric characters are given. Typical localitions are indicated in the map.

Reference:
Gonzalez Sponga MA. Biodiversidad en Venezuela. Descripcion de cuatro nuevas especies del genero Tityus Koch, 1836 (Scorpionida: Buthidae) de los estados Monagas, Sucre y Bolivar. Bol Acad C Fis Mat y Nat. 2008;68 (4):9-30.

Family Buthidae